Millerella yowarensis zone to Pseudostaffella antiqua zone
Upper Serpukhovian to middle Bashkirian.
Microbes/algae always dominating over chaetetids. Maximum chaetetid cover is 27%. Microbes 50-72%, corals max. 8%.
Stratigraphic range of chaetetids from Nagatophyllym satoi Zone (upper Visean) to the Beedina akizoshiensis Zone (upper Moscovian).